SlideShare a Scribd company logo
1 of 23
 What is Biodiversity
 Importance of Biodiversity
 Levels of Biodiversity
 Threats to Biodiversity
 Patterns of Biodiversity
What is Biodiversity?
 The variety of different types of organisms present
and interacting in an ecosystem.
 Often more species equals more diversity, although
there are, in fact many more factors beyond a simple
count of species that determine whether biodiversity
is higher or lower in any given ecosystem.
Biodiversity and global economy
 Globally agriculture, which depends on genetic stock from
natural ecological systems, is now a $3 trillion global
 Recreation and nature tourism generates some $12 billion
worldwide in annual revenues
 In the United States, the economic benefits from wild plants
and animals comprise approximately 4.5% of the Gross
Domestic Product.
 Global trade in wild plants (timber and others) is estimated at
$6 billion annually
Biodiversity and food security
 Much of the world's major food crops, including corn, wheat,
and soybeans, depend on new genetic material from the wild to
remain productive and healthy.
 Food production from wild stocks of fish is the single largest
source of animal protein for the world's 6 billion inhabitants. In
the US alone more than 10 billion pounds of fish, valued at
about $4 billion, were caught and sold yearly.
Levels of Biodiversity
 Genetic Diversity
 Species Diversity
 Ecosystem Diversity
Genetic Diversity
 Amount and variety of genetic material within individuals,
populations or communities
 Source of biodiversity at all levels
 Knowledge of amount of genetic variability present within local
populations essential in directing conservation programs.
 Amount of genetic differences among species could help
determine rates of evolutionary change
Species Level
 Species Richness: numerical count of species present in an
area. Richness tends to increase over area and sampling
intensity
 Species Diversity: When species are weighted by some
measure of importance e.g. abundance, productivity or size.
 Measures of Diversity include:
– Shannon-Wiener Index
– Simpson index
Shannon’s Diversity Index
 Assume that there are n possible categories in a data set and
that their proportions are pi,.....,pn. Then Shannon’s diversity
index for this system is defined to be :
 H’ = -Σpiln(pi)
 accounts for both abundance and evenness of the species
present
 The proportion of species i relative to the total number of
species (pi) is calculated, and then multiplied by the natural
logarithm of this proportion (lnpi).
Simpson’s Diversity Index, D
 Simpson's diversity index (D) characterizes species
diversity in a community.
 Simpson's diversity index (D) characterizes species
diversity in a community.
 D = 1/(Σpi
2
)
 The proportion of species i relative to the total
number of species (pi) is calculated and squared.
The squared proportions for all the species are
summed, and the reciprocal is taken.
Ecosystem and Biome diversity
 Ecosystems are the collection of all the plants and
animals within a particular area
 Ecosystems may differ in species composition,
physical structure and function as a result of
differences in physical structure and composition
 Biomes are large ecological units on the basis of
dominant vegetation
 Preserving a variety of ecosystems and biomes are
necessary for preserving species diversity
Temporal Patterns of Species richness
 Fossil record indicate variation of species
richness over time and space
 Largest number of phyla in the Cambrian
and pre-Cambrian period
 Total number of phyla has since declined but
overall richness has increased
Spatial patterns of species richness
 Point Richness: number of species that can be
found in a single point in space
 Alpha (α-) richness: number of species found in a
small homogenous area
 Beta (β-) richness: rate of change in species in
species composition across habitats
 Gamma (γ-) richness: change across large
landscape gradients
 Richness is directly related to physical environment,
productivity and structural complexity of communities
10000010 100 1000 10000
10
100
Redonda
Saba
Monserrat
Jamaica
Cuba
Area (sq.mi)
Numberofspecies
Relationship between area and number of amphibian
species in selected Islands in West Indies- MacArthur & Wilson 1967
Species /Area relations
Limits of species richness
 Productivity hypothesis: High productivity
results in higher number of species
 Stability hypothesis- environments that are
stable tend to support higher number species
Threats to biodiversity
 habitat destruction (slash and burn agric. or felling
of old-growth forests)
 overexploitation (fishing, hunting)
 pollution (domestic and industrial emissions)
 global climate change (the greenhouse effect and
destruction of the ozone layer)
 invasion by introduced species (displacement of
native species
 underlying social conditions (increased per-capita
consumption, poverty, rapid population growth,
unsound economic and social policies )
Threats to Biodiversity cont’d
 Habitat degradation
– Some 93% of coral reefs damaged directly or
indirectly by human activities
– During the 1990s between 130,000 and 150,000
km2 of forest cover lost each year
 Changes in atmospheric composition.
 siltation, nutrient loading, pollution of air and
water by toxic chemicals
Patterns of species vulnerability
 Rare Species
 Long-lived species
 Keystone species
Rare species
 May be the result of many factors small
range, high habitat specificity or small
population density
 Human-induced rarity may be more
damaging
Long-lived species
 Well-suited to long-term predictability
 Often not equipped to adapt to rapid changes
brought by human-induced changes
 Often population declines may take many
years to recover
Keystone species
 A species or group of species that makes
and unusual contribution to a community
structure or processes
 May be predators, food source or species
that maintains critical ecosystem processes
 A loss of a keystone species may lead to
loss of others that depend on it.
Biodiversity Management
Conservation vs Preservation?
 All about management of Genetic Variation
– Aim is to allow continued evolutionary change in the
populations and species concerned
– Since ecological systems are not static- management
should allow for change- Conservation rather than
preservation.
– 3 Time scales of concern: extinction avoidance (short-term);
ability to adapt or evolve (medium term) and potential for
continued speciation (long-term)
– Units of conservation: What are the units of conservation?
How do we determine the most appropriate unit?
Next week
 Habitat fragmentation and biological
consequences
 Population dynamics on heterogeneous
landscapes
Today’s lab
 Review of two short papers.
 Stuart Chapin III et al 2000. Consequences of changing
biodiversity Nature Vol. 405 pp. 234 http://www.nature.com/cgi-
taf/DynaPage.taf?file=/nature/journal/v405/n6783/full/405234a0_fs.html&content_filetype=pdf
 Franklin, J.F. 1993. Preserving Biodiversity: Species,
Ecosystems or Landscapes? Ecological Applications, 3(2), pp.
202 - 205. http://www.jstor.org/cgi-bin/jstor/printpage/10510761/di960380/96p0004u/0.pd f?
userID=a027019f@columbia.edu/01cc9933410050dc70eb&backcontext=table-
ofcontents&config=jstor&dowhat=Acrobat&0.pdf

More Related Content

What's hot

Introduction To Ecology
Introduction To EcologyIntroduction To Ecology
Introduction To Ecology
sacklax40
 

What's hot (20)

Importance of biodiversity to humanKinds
Importance of biodiversity to humanKindsImportance of biodiversity to humanKinds
Importance of biodiversity to humanKinds
 
Biodiversity conservation strategies
Biodiversity conservation strategiesBiodiversity conservation strategies
Biodiversity conservation strategies
 
Biodiversity
BiodiversityBiodiversity
Biodiversity
 
Introduction To Ecology
Introduction To EcologyIntroduction To Ecology
Introduction To Ecology
 
Ppt on biodiversity
Ppt on biodiversityPpt on biodiversity
Ppt on biodiversity
 
Biodiversity
BiodiversityBiodiversity
Biodiversity
 
Conservation biology note pdf
Conservation biology note pdfConservation biology note pdf
Conservation biology note pdf
 
Presentation of biodiversity
Presentation of biodiversityPresentation of biodiversity
Presentation of biodiversity
 
Biodiversity
BiodiversityBiodiversity
Biodiversity
 
Community ecology
Community ecology Community ecology
Community ecology
 
Threats to biological diversity
Threats to biological diversityThreats to biological diversity
Threats to biological diversity
 
Loss of biodiversity
Loss of biodiversityLoss of biodiversity
Loss of biodiversity
 
Distribution in global pattern of biodiversity
Distribution in global pattern of biodiversityDistribution in global pattern of biodiversity
Distribution in global pattern of biodiversity
 
Lesson 1. concept of ecology
Lesson 1. concept of ecologyLesson 1. concept of ecology
Lesson 1. concept of ecology
 
Biodiversity and its conservation
Biodiversity and its conservationBiodiversity and its conservation
Biodiversity and its conservation
 
Ecological efficiency
Ecological efficiencyEcological efficiency
Ecological efficiency
 
best ever ppt on ecological succession by nagesh
best ever ppt on  ecological succession by nageshbest ever ppt on  ecological succession by nagesh
best ever ppt on ecological succession by nagesh
 
Metapopulation
MetapopulationMetapopulation
Metapopulation
 
Ecoregions
EcoregionsEcoregions
Ecoregions
 
Ecological succession
Ecological successionEcological succession
Ecological succession
 

Similar to Biodiversity ist lecture

Populations
PopulationsPopulations
Populations
Elisa
 
Answer (6)THREE WAYS HUMAN HAVE NEGATIVELY IMPACTED BIODIVERSITY .pdf
Answer (6)THREE WAYS HUMAN HAVE NEGATIVELY IMPACTED BIODIVERSITY .pdfAnswer (6)THREE WAYS HUMAN HAVE NEGATIVELY IMPACTED BIODIVERSITY .pdf
Answer (6)THREE WAYS HUMAN HAVE NEGATIVELY IMPACTED BIODIVERSITY .pdf
angelsfashion1
 
Deocareza populationecology
Deocareza populationecologyDeocareza populationecology
Deocareza populationecology
carlo2307
 
Deocareza population ecology-1231427563650176-1 (1)
Deocareza population ecology-1231427563650176-1 (1)Deocareza population ecology-1231427563650176-1 (1)
Deocareza population ecology-1231427563650176-1 (1)
carlo2307
 
Population ecology
Population ecologyPopulation ecology
Population ecology
Crystal Wood
 

Similar to Biodiversity ist lecture (20)

Lecture note on Biodiversity conservation
Lecture note on Biodiversity conservationLecture note on Biodiversity conservation
Lecture note on Biodiversity conservation
 
PopulationsChapter_9-10.ppt
PopulationsChapter_9-10.pptPopulationsChapter_9-10.ppt
PopulationsChapter_9-10.ppt
 
Populations
PopulationsPopulations
Populations
 
Populations
PopulationsPopulations
Populations
 
BIODIVERSITY & ITS TYPES
BIODIVERSITY & ITS TYPESBIODIVERSITY & ITS TYPES
BIODIVERSITY & ITS TYPES
 
BIOLOGICAL DIVERSITY
BIOLOGICAL DIVERSITYBIOLOGICAL DIVERSITY
BIOLOGICAL DIVERSITY
 
2.1 Population Dynamics new revision slides
2.1  Population Dynamics new revision slides2.1  Population Dynamics new revision slides
2.1 Population Dynamics new revision slides
 
Biodiversity
BiodiversityBiodiversity
Biodiversity
 
for LIVE Teaching Q3 Week 4- Ecosystem.pptx
for LIVE Teaching Q3 Week 4- Ecosystem.pptxfor LIVE Teaching Q3 Week 4- Ecosystem.pptx
for LIVE Teaching Q3 Week 4- Ecosystem.pptx
 
for LIVE Teaching Q3 Week 4- Ecosystem.pptx
for LIVE Teaching Q3 Week 4- Ecosystem.pptxfor LIVE Teaching Q3 Week 4- Ecosystem.pptx
for LIVE Teaching Q3 Week 4- Ecosystem.pptx
 
Ch 6 ed
Ch 6 edCh 6 ed
Ch 6 ed
 
Population growth
Population growthPopulation growth
Population growth
 
Answer (6)THREE WAYS HUMAN HAVE NEGATIVELY IMPACTED BIODIVERSITY .pdf
Answer (6)THREE WAYS HUMAN HAVE NEGATIVELY IMPACTED BIODIVERSITY .pdfAnswer (6)THREE WAYS HUMAN HAVE NEGATIVELY IMPACTED BIODIVERSITY .pdf
Answer (6)THREE WAYS HUMAN HAVE NEGATIVELY IMPACTED BIODIVERSITY .pdf
 
Essay Biodiversity
Essay BiodiversityEssay Biodiversity
Essay Biodiversity
 
LEVELS OF BIO-DIVERSITY 52,53,54 final (1).pptx
LEVELS OF BIO-DIVERSITY 52,53,54 final (1).pptxLEVELS OF BIO-DIVERSITY 52,53,54 final (1).pptx
LEVELS OF BIO-DIVERSITY 52,53,54 final (1).pptx
 
Lecture 11 biodiversity
Lecture 11 biodiversityLecture 11 biodiversity
Lecture 11 biodiversity
 
Zoology Seminar 2011
Zoology Seminar 2011Zoology Seminar 2011
Zoology Seminar 2011
 
Deocareza populationecology
Deocareza populationecologyDeocareza populationecology
Deocareza populationecology
 
Deocareza population ecology-1231427563650176-1 (1)
Deocareza population ecology-1231427563650176-1 (1)Deocareza population ecology-1231427563650176-1 (1)
Deocareza population ecology-1231427563650176-1 (1)
 
Population ecology
Population ecologyPopulation ecology
Population ecology
 

Recently uploaded

MSc Ag Genetics & Plant Breeding: Insights from Previous Year JNKVV Entrance ...
MSc Ag Genetics & Plant Breeding: Insights from Previous Year JNKVV Entrance ...MSc Ag Genetics & Plant Breeding: Insights from Previous Year JNKVV Entrance ...
MSc Ag Genetics & Plant Breeding: Insights from Previous Year JNKVV Entrance ...
Krashi Coaching
 
SURVEY I created for uni project research
SURVEY I created for uni project researchSURVEY I created for uni project research
SURVEY I created for uni project research
CaitlinCummins3
 
會考英文會考英文會考英文會考英文會考英文會考英文會考英文會考英文會考英文會考英文會考英文
會考英文會考英文會考英文會考英文會考英文會考英文會考英文會考英文會考英文會考英文會考英文會考英文會考英文會考英文會考英文會考英文會考英文會考英文會考英文會考英文會考英文會考英文
會考英文會考英文會考英文會考英文會考英文會考英文會考英文會考英文會考英文會考英文會考英文
中 央社
 

Recently uploaded (20)

Word Stress rules esl .pptx
Word Stress rules esl               .pptxWord Stress rules esl               .pptx
Word Stress rules esl .pptx
 
size separation d pharm 1st year pharmaceutics
size separation d pharm 1st year pharmaceuticssize separation d pharm 1st year pharmaceutics
size separation d pharm 1st year pharmaceutics
 
Incoming and Outgoing Shipments in 2 STEPS Using Odoo 17
Incoming and Outgoing Shipments in 2 STEPS Using Odoo 17Incoming and Outgoing Shipments in 2 STEPS Using Odoo 17
Incoming and Outgoing Shipments in 2 STEPS Using Odoo 17
 
Mattingly "AI and Prompt Design: LLMs with Text Classification and Open Source"
Mattingly "AI and Prompt Design: LLMs with Text Classification and Open Source"Mattingly "AI and Prompt Design: LLMs with Text Classification and Open Source"
Mattingly "AI and Prompt Design: LLMs with Text Classification and Open Source"
 
An Overview of the Odoo 17 Knowledge App
An Overview of the Odoo 17 Knowledge AppAn Overview of the Odoo 17 Knowledge App
An Overview of the Odoo 17 Knowledge App
 
Software testing for project report .pdf
Software testing for project report .pdfSoftware testing for project report .pdf
Software testing for project report .pdf
 
Graduate Outcomes Presentation Slides - English (v3).pptx
Graduate Outcomes Presentation Slides - English (v3).pptxGraduate Outcomes Presentation Slides - English (v3).pptx
Graduate Outcomes Presentation Slides - English (v3).pptx
 
MSc Ag Genetics & Plant Breeding: Insights from Previous Year JNKVV Entrance ...
MSc Ag Genetics & Plant Breeding: Insights from Previous Year JNKVV Entrance ...MSc Ag Genetics & Plant Breeding: Insights from Previous Year JNKVV Entrance ...
MSc Ag Genetics & Plant Breeding: Insights from Previous Year JNKVV Entrance ...
 
MOOD STABLIZERS DRUGS.pptx
MOOD     STABLIZERS           DRUGS.pptxMOOD     STABLIZERS           DRUGS.pptx
MOOD STABLIZERS DRUGS.pptx
 
An overview of the various scriptures in Hinduism
An overview of the various scriptures in HinduismAn overview of the various scriptures in Hinduism
An overview of the various scriptures in Hinduism
 
MichaelStarkes_UncutGemsProjectSummary.pdf
MichaelStarkes_UncutGemsProjectSummary.pdfMichaelStarkes_UncutGemsProjectSummary.pdf
MichaelStarkes_UncutGemsProjectSummary.pdf
 
“O BEIJO” EM ARTE .
“O BEIJO” EM ARTE                       .“O BEIJO” EM ARTE                       .
“O BEIJO” EM ARTE .
 
Exploring Gemini AI and Integration with MuleSoft | MuleSoft Mysore Meetup #45
Exploring Gemini AI and Integration with MuleSoft | MuleSoft Mysore Meetup #45Exploring Gemini AI and Integration with MuleSoft | MuleSoft Mysore Meetup #45
Exploring Gemini AI and Integration with MuleSoft | MuleSoft Mysore Meetup #45
 
SURVEY I created for uni project research
SURVEY I created for uni project researchSURVEY I created for uni project research
SURVEY I created for uni project research
 
會考英文會考英文會考英文會考英文會考英文會考英文會考英文會考英文會考英文會考英文會考英文
會考英文會考英文會考英文會考英文會考英文會考英文會考英文會考英文會考英文會考英文會考英文會考英文會考英文會考英文會考英文會考英文會考英文會考英文會考英文會考英文會考英文會考英文
會考英文會考英文會考英文會考英文會考英文會考英文會考英文會考英文會考英文會考英文會考英文
 
ANTI PARKISON DRUGS.pptx
ANTI         PARKISON          DRUGS.pptxANTI         PARKISON          DRUGS.pptx
ANTI PARKISON DRUGS.pptx
 
Dementia (Alzheimer & vasular dementia).
Dementia (Alzheimer & vasular dementia).Dementia (Alzheimer & vasular dementia).
Dementia (Alzheimer & vasular dementia).
 
UChicago CMSC 23320 - The Best Commit Messages of 2024
UChicago CMSC 23320 - The Best Commit Messages of 2024UChicago CMSC 23320 - The Best Commit Messages of 2024
UChicago CMSC 23320 - The Best Commit Messages of 2024
 
II BIOSENSOR PRINCIPLE APPLICATIONS AND WORKING II
II BIOSENSOR PRINCIPLE APPLICATIONS AND WORKING IIII BIOSENSOR PRINCIPLE APPLICATIONS AND WORKING II
II BIOSENSOR PRINCIPLE APPLICATIONS AND WORKING II
 
philosophy and it's principles based on the life
philosophy and it's principles based on the lifephilosophy and it's principles based on the life
philosophy and it's principles based on the life
 

Biodiversity ist lecture

  • 1.  What is Biodiversity  Importance of Biodiversity  Levels of Biodiversity  Threats to Biodiversity  Patterns of Biodiversity
  • 2. What is Biodiversity?  The variety of different types of organisms present and interacting in an ecosystem.  Often more species equals more diversity, although there are, in fact many more factors beyond a simple count of species that determine whether biodiversity is higher or lower in any given ecosystem.
  • 3. Biodiversity and global economy  Globally agriculture, which depends on genetic stock from natural ecological systems, is now a $3 trillion global  Recreation and nature tourism generates some $12 billion worldwide in annual revenues  In the United States, the economic benefits from wild plants and animals comprise approximately 4.5% of the Gross Domestic Product.  Global trade in wild plants (timber and others) is estimated at $6 billion annually
  • 4. Biodiversity and food security  Much of the world's major food crops, including corn, wheat, and soybeans, depend on new genetic material from the wild to remain productive and healthy.  Food production from wild stocks of fish is the single largest source of animal protein for the world's 6 billion inhabitants. In the US alone more than 10 billion pounds of fish, valued at about $4 billion, were caught and sold yearly.
  • 5. Levels of Biodiversity  Genetic Diversity  Species Diversity  Ecosystem Diversity
  • 6. Genetic Diversity  Amount and variety of genetic material within individuals, populations or communities  Source of biodiversity at all levels  Knowledge of amount of genetic variability present within local populations essential in directing conservation programs.  Amount of genetic differences among species could help determine rates of evolutionary change
  • 7. Species Level  Species Richness: numerical count of species present in an area. Richness tends to increase over area and sampling intensity  Species Diversity: When species are weighted by some measure of importance e.g. abundance, productivity or size.  Measures of Diversity include: – Shannon-Wiener Index – Simpson index
  • 8. Shannon’s Diversity Index  Assume that there are n possible categories in a data set and that their proportions are pi,.....,pn. Then Shannon’s diversity index for this system is defined to be :  H’ = -Σpiln(pi)  accounts for both abundance and evenness of the species present  The proportion of species i relative to the total number of species (pi) is calculated, and then multiplied by the natural logarithm of this proportion (lnpi).
  • 9. Simpson’s Diversity Index, D  Simpson's diversity index (D) characterizes species diversity in a community.  Simpson's diversity index (D) characterizes species diversity in a community.  D = 1/(Σpi 2 )  The proportion of species i relative to the total number of species (pi) is calculated and squared. The squared proportions for all the species are summed, and the reciprocal is taken.
  • 10. Ecosystem and Biome diversity  Ecosystems are the collection of all the plants and animals within a particular area  Ecosystems may differ in species composition, physical structure and function as a result of differences in physical structure and composition  Biomes are large ecological units on the basis of dominant vegetation  Preserving a variety of ecosystems and biomes are necessary for preserving species diversity
  • 11. Temporal Patterns of Species richness  Fossil record indicate variation of species richness over time and space  Largest number of phyla in the Cambrian and pre-Cambrian period  Total number of phyla has since declined but overall richness has increased
  • 12. Spatial patterns of species richness  Point Richness: number of species that can be found in a single point in space  Alpha (α-) richness: number of species found in a small homogenous area  Beta (β-) richness: rate of change in species in species composition across habitats  Gamma (γ-) richness: change across large landscape gradients  Richness is directly related to physical environment, productivity and structural complexity of communities
  • 13. 10000010 100 1000 10000 10 100 Redonda Saba Monserrat Jamaica Cuba Area (sq.mi) Numberofspecies Relationship between area and number of amphibian species in selected Islands in West Indies- MacArthur & Wilson 1967 Species /Area relations
  • 14. Limits of species richness  Productivity hypothesis: High productivity results in higher number of species  Stability hypothesis- environments that are stable tend to support higher number species
  • 15. Threats to biodiversity  habitat destruction (slash and burn agric. or felling of old-growth forests)  overexploitation (fishing, hunting)  pollution (domestic and industrial emissions)  global climate change (the greenhouse effect and destruction of the ozone layer)  invasion by introduced species (displacement of native species  underlying social conditions (increased per-capita consumption, poverty, rapid population growth, unsound economic and social policies )
  • 16. Threats to Biodiversity cont’d  Habitat degradation – Some 93% of coral reefs damaged directly or indirectly by human activities – During the 1990s between 130,000 and 150,000 km2 of forest cover lost each year  Changes in atmospheric composition.  siltation, nutrient loading, pollution of air and water by toxic chemicals
  • 17. Patterns of species vulnerability  Rare Species  Long-lived species  Keystone species
  • 18. Rare species  May be the result of many factors small range, high habitat specificity or small population density  Human-induced rarity may be more damaging
  • 19. Long-lived species  Well-suited to long-term predictability  Often not equipped to adapt to rapid changes brought by human-induced changes  Often population declines may take many years to recover
  • 20. Keystone species  A species or group of species that makes and unusual contribution to a community structure or processes  May be predators, food source or species that maintains critical ecosystem processes  A loss of a keystone species may lead to loss of others that depend on it.
  • 21. Biodiversity Management Conservation vs Preservation?  All about management of Genetic Variation – Aim is to allow continued evolutionary change in the populations and species concerned – Since ecological systems are not static- management should allow for change- Conservation rather than preservation. – 3 Time scales of concern: extinction avoidance (short-term); ability to adapt or evolve (medium term) and potential for continued speciation (long-term) – Units of conservation: What are the units of conservation? How do we determine the most appropriate unit?
  • 22. Next week  Habitat fragmentation and biological consequences  Population dynamics on heterogeneous landscapes
  • 23. Today’s lab  Review of two short papers.  Stuart Chapin III et al 2000. Consequences of changing biodiversity Nature Vol. 405 pp. 234 http://www.nature.com/cgi- taf/DynaPage.taf?file=/nature/journal/v405/n6783/full/405234a0_fs.html&content_filetype=pdf  Franklin, J.F. 1993. Preserving Biodiversity: Species, Ecosystems or Landscapes? Ecological Applications, 3(2), pp. 202 - 205. http://www.jstor.org/cgi-bin/jstor/printpage/10510761/di960380/96p0004u/0.pd f? userID=a027019f@columbia.edu/01cc9933410050dc70eb&backcontext=table- ofcontents&config=jstor&dowhat=Acrobat&0.pdf

Editor's Notes

  1. World Resources Institute, The World Conservation Union, and United Nations Environment Program. 1992. Global Biodiversity Strategy. Washington, D.C.: World Resources Institute World Conservation Monitoring Center (WCMC). 1992. Global Biodiversity: Status of the Earth's Living Resources. Redding, U.K.: WCMC.
  2. National Oceanic and Atmospheric Administration (NOAA). 1995. Fisheries of the United States, 1994. Washington, D.C.: U.S. Government Printing Office.
  3. Shannon-Wiener index H’ = Sum(pi)Ln(pi) Simpson index D = 1/(sum (pi2)