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The mythos of the researcher
Monitoring – Long-term Value
• The information never goes out of date
• The information NOT collected now can NOT be
  collected later
• I and M data are used many times by many
  people
• I and M data creates the foundation for Research
  and Researchers
• The names and legacies of these people are kept
  in our memories
• Many are NOT professionals
Hawaii Pacific GIS Conference 2012: GIS for Citizen Engagement - Hawaii Coqui Crawl Project - Counting Frogs with Cell Phones
How do we know there are Coqui
        Frogs in Hawaii?
• Native Crickets?
• Introduced Katydids?
• Etc.?
Counting Crickets with Cell
Phones: How we Harnessed
 New Yorkers’ Favorite Tool
During the NYC Cricket Crawl

              Lou Sorkin, Sam Droege,
                 Elizabeth Johnson,
           John Pickering, Tammy Pittman,
                     Emily Sweet
Goals

• Collect scientifically defensible data about
  “singing” Orthoptera in metro region
  – Survey 7 easy to ID species
  – Look for Common True Katydid colonies
     • William T. Davis mentions them being absent [The True
       Katydid nearly extinct in New York City] in the early part of
       last century, published in the Journal of the New York
       Entomological Society. Vol. 28(1), March 1920)
More Goals
• Engage the public in data collection
  – have scientific, non-scientific groups, and citizens work
    together
  – generate excitement and intrigue to lure people into going
    outside, thinking about nature, and possibly pursuing
    natural history study
• Test out new citizen science protocols
  – people sending in data immediately from the field
  – entering and displaying data by the end of the night
More, More Goals
• Integrate art into a scientific venture
• Set an example for what other groups could
  replicate elsewhere
• Do it for 0 dollars
• Have fun
Quick Overview
• September 12, 2009 – rain date
• Advertised via list serves, facebook, emails,
  other PR
• Data quality
  – selected 7 relatively easy to ID species
  – set up website with training materials, audio files
  – encouraged group survey (allowing folks to confer
    on ID)
  – expert field confirmation of questionable
    submissions
Participation
• ~ 400 sites surveyed
• ~ 300 individuals involved
  – Most people went out in small groups
     •   Families went home afterwards
     •   Other people went out to bars afterwards
     •   Some worked overnight into the morning
     •   Some ??
Data Submission
• Information submitted:
  – Observer name, exact location, time, species
    heard
• Phone message (converted to .mp3file)
  or email linked to drop.io box and picked
  up by workers at Cricket Crawl HQ
• Data entered by people at Cricket Crawl HQ
  and on the Internet – listen, transcribe
Cricket Crawl Headquarters
Maps of the Results
• Generated by John Pickering at the University
  of Georgia
• Programmed to update every 10 minutes
  throughout the night
Map Legend

• Yellow dots are sampling locations

• Red dots are places that recorded that species
Orocharis saltator
           Jumping Bush Cricket

• Common species
• Potentially some confusion with Snowy Tree
  Crickets (not included in the survey) and with
  Fall Field Crickets
• Not mentioned by Davis as present during his
  time
Orocharis saltator
Jumping Bush Cricket
Orocharis saltator
Jumping Bush Cricket
Example of Drilling Down to
Manhattan Using the Online Mapping
               Tool
Drilling Further Down
And Further Down
Clicking on a Dot Shows you the Data and if
  there was a phone call you can play it to
             hear the observer
Gryllus pennsylvanicus
          Fall Field Cricket
• Common and ubiquitous
• Ground-dwelling
Gryllus pennsylvanicus
   Fall Field Cricket
Gryllus pennsylvanicus
   Fall Field Cricket
Pterophylla camellifolia
         Common True Katydid
• Large clumsy species that appears in small colonies
• Essentially doesn’t fly
• Disappeared from Staten Island in the early part of the
  last century
• Only reappeared lately (re: Paul Lederer and co. work)
• Unclear how long present on Manhattan
• Distribution clumped, but present in some very small
  woodlots (re: Marie Winn’s)
• Follow-up to confirm some localities
   needed
Pterophylla camellifolia
Common True Katydid
Pterophylla camellifolia
 Common true katydid
Gowanus Expedition
Things People Liked ….

• Being outside at night looking at something
  they thought they knew, but really didn’t
• Sending in data with their cell phone
• Being part of an event
• Learning about crickets and katydids in general
•   Dociostaurus (Dociostaurus) maroccanus (Thunberg, 1815)   •   Prognathogryllus inexspectatus Perkins, 1899                •   Trigonidium mahina Otte, 1994
•   Gomphocerus plebejus Stål, 1861                           •   Prognathogryllus kahea Otte, 1994                           •   Trigonidium makani Otte, 1994
•   Gryllidae                                                 •   Prognathogryllus kahili Otte, 1994                          •   Trigonidium makanina Otte, 1994
•   Anele ulia Otte, Carvalho & Shaw, 2003                    •   Prognathogryllus kipahulu Otte, 1994                        •   Trigonidium makapala Otte, 1994
•   Caconemobius albus Otte, 1994                             •   Prognathogryllus kohala Otte, 1994                          •   Trigonidium makau Otte, 1994
•   Caconemobius anahulu Otte, 1994                           •   Prognathogryllus kukui Otte, 1994                           •   Trigonidium malanai Otte, 1994
•   Caconemobius fori Gurney & Rentz, 1978                    •   Prognathogryllus makai Otte, 1994                           •   Trigonidium mana Otte, 1994
•   Caconemobius howarthi Gurney & Rentz, 1978                •   Prognathogryllus makakapua Otte, 1994                       •   Trigonidium manuka Carvalho & Otte, 2006
•   Caconemobius nihoensis Otte, 1994                         •   Prognathogryllus mauka Otte, 1994                           •   Trigonidium mauiensis Otte, 1994
•   Caconemobius paralbus Otte, 1994                          •   Prognathogryllus oahuensis Perkins, 1899                    •   Trigonidium mauka Otte, 1994
•   Caconemobius sandwichensis Otte, 1994                     •   Prognathogryllus olympus Otte, 1994                         •   Trigonidium mokuleia Carvalho & Otte, 2006
•   Caconemobius uuku Otte, 1994                              •   Prognathogryllus opua Otte, 1994                            •   Trigonidium molokaiense (Perkins, 1899)
•   Caconemobius varius Gurney & Rentz, 1978                  •   Prognathogryllus parakahili Otte, 1994                      •   Trigonidium nani Otte, 1994
•   Laupala cerasina Otte, 1994                               •   Prognathogryllus parakukui Otte, 1994                       •   Trigonidium napua Carvalho & Otte, 2006
•   Laupala eukolea Otte, 1994                                •   Prognathogryllus pararobustus Otte, 1994                    •   Trigonidium nele Otte, 1994
•   Laupala eupacifica Otte, 1994                             •   Prognathogryllus pihea Otte, 1994                           •   Trigonidium neogrande Otte, 1994
•   Laupala fugax Otte, 1994                                  •   Prognathogryllus puna Otte, 1994                            •   Trigonidium neokukui Otte, 1994
•   Laupala hapapa Otte, 1994                                 •   Prognathogryllus robustus Perkins, 1899                     •   Trigonidium neovarians Otte, 1994
•   Laupala hualalai Otte, 1994                               •   Prognathogryllus spadix Otte, 1994                          •   Trigonidium novagintum Otte, 1994
•   Laupala kai Otte, 1994                                    •   Prognathogryllus stridulans (Perkins, 1899)                 •   Trigonidium novena Otte, 1994
•   Laupala kanaele Otte, 1994                                •   Prognathogryllus victoriae Otte, 1994                       •   Trigonidium octavum Otte, 1994
•   Laupala kauaiensis Otte, 1994                             •   Prognathogryllus waikemoi Otte, 1994                        •   Trigonidium octonalis Otte, 1994
•   Laupala kohalensis Otte, 1994                             •   Prognathogryllus weli Otte, 1994                            •   Trigonidium ohaka Otte, 1994
•   Laupala kokeensis Otte, 1994                              •   Prolaupala kaalensis Otte, 1994                             •   Trigonidium ola Otte, 1994
•   Laupala kolea Otte, 1994                                  •   Prolaupala kukui Otte, 1994                                 •   Trigonidium olomea Otte, 1994
•   Laupala koloa Otte, 1994                                  •   Prolaupala perkinsi Otte, 1994                              •   Trigonidium ookala Otte, 1994
•   Laupala kona Otte, 1994                                   •   Thaumatogryllus cavicola Gurney & Rentz, 1978               •   Trigonidium opua Otte, 1994
•   Laupala lanaiensis Otte, 1994                             •   Thaumatogryllus conanti Otte, 1994                          •   Trigonidium pahiwa Otte, 1994
•   Laupala makaio Shaw, 2000                                 •   Thaumatogryllus mauiensis Otte, 1994                        •   Trigonidium palai Otte, 1994
•   Laupala makaweli Otte, 1994                               •   Thaumatogryllus variegatus Perkins, 1899                    •   Trigonidium paramana Otte, 1994
•   Laupala media Otte, 1994                                  •   Trigonidium acuste Otte, 1994                               •   Trigonidium paranoe Otte, 1994
•   Laupala mediaspisa Otte, 1994                             •   Trigonidium ahiu Otte, 1994                                 •   Trigonidium paranoho Otte, 1994
•   Laupala melewiki Shaw, 2000                               •   Trigonidium aka Otte, 1994                                  •   Trigonidium paraspilos Otte, 1994
•   Laupala molokaiensis Otte, 1994                           •   Trigonidium akaka Otte, 1994                                •   Trigonidium paroctonalis Otte, 1994
•   Laupala neospisa Otte, 1994                               •   Trigonidium alina Otte, 1994                                •   Trigonidium pavida Otte, 1994
•   Laupala nigra Otte, 1994                                  •   Trigonidium alternatum (Perkins, 1899)                      •   Trigonidium pipili Otte, 1994
•   Laupala nui Otte, 1994                                    •   Trigonidium anoe Otte, 1994                                 •   Trigonidium pololu Otte, 1994
•   Laupala oahuensis Otte, 1994                              •   Trigonidium atroferrugineum (Brunner von Wattenwyl, 1895)   •   Trigonidium proalina Otte, 1994
•   Laupala olohena Otte, 1994                                •   Trigonidium attenuatum (Perkins, 1899)                      •   Trigonidium procrustum Otte, 1994
•   Laupala pacifica Otte, 1994                               •   Trigonidium awawa Otte, 1994                                •   Trigonidium promana Otte, 1994
•   Laupala paranigra Otte, 1994                              •   Trigonidium awiwi Otte, 1994                                •   Trigonidium pseudokua Otte, 1994
•   Laupala parapacifica Otte, 1994                           •   Trigonidium crepitans (Perkins, 1899)                       •   Trigonidium pseudoli Otte, 1994
•   Laupala paraprosea Otte, 1994                             •   Trigonidium debile (Perkins, 1899)                          •   Trigonidium pseudonoe Otte, 1994
•   Laupala prosea Otte, 1994                                 •   Trigonidium eumeles Otte, 1994                              •   Trigonidium pudicum Otte, 1994
•   Laupala pruna Otte, 1994                                  •   Trigonidium excultatum Otte, 1994                           •   Trigonidium puiwa Otte, 1994
•   Laupala spisa Otte, 1994                                  •   Trigonidium exiguum Perkins, 1899                           •   Trigonidium puukani Otte, 1994
•   Laupala tantalis Otte, 1994                               •   Trigonidium exuberans Otte, 1994                            •   Trigonidium robustum Perkins, 1899
•   Laupala vespertina Otte, 1994                             •   Trigonidium filicum (Perkins, 1899)                         •   Trigonidium roseum (Perkins, 1899)
•   Laupala waikemoi Otte, 1994                               •   Trigonidium flectens Otte, 1994                             •   Trigonidium saltator (Perkins, 1899)
•   Laupala wailua Otte, 1994                                 •   Trigonidium fortuitum Otte, 1994                            •   Trigonidium septimum Otte, 1994
•   Leptogryllus apicalis Perkins, 1910                       •   Trigonidium freycinetiae (Perkins, 1899)                    •   Trigonidium sextum Otte, 1994
•   Leptogryllus deceptor Perkins, 1910                       •   Trigonidium fritinnium Otte, 1994                           •   Trigonidium sibilans Otte, 1994
•   Leptogryllus elongatus Perkins, 1899                      •   Trigonidium grande (Perkins, 1899)                          •   Trigonidium silvicola Otte, 1994
•   Leptogryllus forficularis (Brunner von Wattenwyl, 1895)   •   Trigonidium haawina Otte, 1994                              •   Trigonidium spilos Otte, 1994
•   Leptogryllus fusconotatus Perkins, 1899                   •   Trigonidium halulu Otte, 1994                               •   Trigonidium subroseum (Perkins, 1899)
•   Leptogryllus hanaula Otte, 1994                           •   Trigonidium hamakua Otte, 1994                              •   Trigonidium sylvaticum Otte, 1994
•   Leptogryllus haupu Otte, 1994                             •   Trigonidium hamumu Otte, 1994                               •   Trigonidium triens Otte, 1994
•   Leptogryllus kaala Otte, 1994                             •   Trigonidium hapapa Otte, 1994                               •   Trigonidium ua Otte, 1994
•   Leptogryllus kainalu Otte, 1994                           •   Trigonidium haupu Otte, 1994                                •   Trigonidium ulaino Otte, 1994
•   Leptogryllus kauaiensis Perkins, 1899                     •   Trigonidium hehelo Otte, 1994                               •   Trigonidium varians (Perkins, 1899)
•   Leptogryllus kawela Otte, 1994                            •   Trigonidium hoahoa Otte, 1994                               •   Trigonidium venatum Otte, 1994
•   Leptogryllus kipahulu Otte, 1994                          •   Trigonidium hoku Otte, 1994                                 •   Trigonidium virens Otte, 1994
•   Leptogryllus kohala Otte, 1994                            •   Trigonidium holomua Otte, 1994                              •   Trigonidium viridiscens (Perkins, 1899)
•   Leptogryllus lanaiensis Otte, 1994                        •   Trigonidium hopo Otte, 1994                                 •   Trigonidium wahoi Otte, 1994
•   Leptogryllus luteus Otte, 1994                            •   Trigonidium huapala Otte, 1994                              •   Trigonidium wai Otte, 1994
•   Leptogryllus mauiensis Otte, 1994                         •   Trigonidium hyperkona Otte, 1994                            •   Trigonidium waialina Otte, 1994
•   Leptogryllus mauumae Otte, 1994                           •   Trigonidium iao Otte, 1994                                  •   Trigonidium waikua Otte, 1994
•   Leptogryllus molokai Otte, 1994                           •   Trigonidium ignavum Otte, 1994                              •   Trigonidium waimea Otte, 1994
•   Leptogryllus montgomeri Otte, 1994                        •   Trigonidium illex Otte, 1994                                •   Trigonidium waipuna Otte, 1994
•   Leptogryllus nigrolineatus (Perkins, 1899)                •   Trigonidium imitans Otte, 1994                              •   Trigonidium wiki Otte, 1994
•   Leptogryllus nigromaculatus Perkins, 1899                 •   Trigonidium improbum Otte, 1994                             •   Myrmecophilidae
•   Leptogryllus oahuensis Otte, 1994                         •   Trigonidium incongruum Otte, 1994                           •   Myrmecophilus (Myrmecophilus) quadrispina Perkins, 1899
Sam Droege – sdroege@usgs.gov

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Hawaii Pacific GIS Conference 2012: GIS for Citizen Engagement - Hawaii Coqui Crawl Project - Counting Frogs with Cell Phones

  • 1. The mythos of the researcher
  • 2. Monitoring – Long-term Value • The information never goes out of date • The information NOT collected now can NOT be collected later • I and M data are used many times by many people • I and M data creates the foundation for Research and Researchers • The names and legacies of these people are kept in our memories • Many are NOT professionals
  • 4. How do we know there are Coqui Frogs in Hawaii? • Native Crickets? • Introduced Katydids? • Etc.?
  • 5. Counting Crickets with Cell Phones: How we Harnessed New Yorkers’ Favorite Tool During the NYC Cricket Crawl Lou Sorkin, Sam Droege, Elizabeth Johnson, John Pickering, Tammy Pittman, Emily Sweet
  • 6. Goals • Collect scientifically defensible data about “singing” Orthoptera in metro region – Survey 7 easy to ID species – Look for Common True Katydid colonies • William T. Davis mentions them being absent [The True Katydid nearly extinct in New York City] in the early part of last century, published in the Journal of the New York Entomological Society. Vol. 28(1), March 1920)
  • 7. More Goals • Engage the public in data collection – have scientific, non-scientific groups, and citizens work together – generate excitement and intrigue to lure people into going outside, thinking about nature, and possibly pursuing natural history study • Test out new citizen science protocols – people sending in data immediately from the field – entering and displaying data by the end of the night
  • 8. More, More Goals • Integrate art into a scientific venture • Set an example for what other groups could replicate elsewhere • Do it for 0 dollars • Have fun
  • 9. Quick Overview • September 12, 2009 – rain date • Advertised via list serves, facebook, emails, other PR • Data quality – selected 7 relatively easy to ID species – set up website with training materials, audio files – encouraged group survey (allowing folks to confer on ID) – expert field confirmation of questionable submissions
  • 10. Participation • ~ 400 sites surveyed • ~ 300 individuals involved – Most people went out in small groups • Families went home afterwards • Other people went out to bars afterwards • Some worked overnight into the morning • Some ??
  • 11. Data Submission • Information submitted: – Observer name, exact location, time, species heard • Phone message (converted to .mp3file) or email linked to drop.io box and picked up by workers at Cricket Crawl HQ • Data entered by people at Cricket Crawl HQ and on the Internet – listen, transcribe
  • 13. Maps of the Results • Generated by John Pickering at the University of Georgia • Programmed to update every 10 minutes throughout the night
  • 14. Map Legend • Yellow dots are sampling locations • Red dots are places that recorded that species
  • 15. Orocharis saltator Jumping Bush Cricket • Common species • Potentially some confusion with Snowy Tree Crickets (not included in the survey) and with Fall Field Crickets • Not mentioned by Davis as present during his time
  • 18. Example of Drilling Down to Manhattan Using the Online Mapping Tool
  • 21. Clicking on a Dot Shows you the Data and if there was a phone call you can play it to hear the observer
  • 22. Gryllus pennsylvanicus Fall Field Cricket • Common and ubiquitous • Ground-dwelling
  • 23. Gryllus pennsylvanicus Fall Field Cricket
  • 24. Gryllus pennsylvanicus Fall Field Cricket
  • 25. Pterophylla camellifolia Common True Katydid • Large clumsy species that appears in small colonies • Essentially doesn’t fly • Disappeared from Staten Island in the early part of the last century • Only reappeared lately (re: Paul Lederer and co. work) • Unclear how long present on Manhattan • Distribution clumped, but present in some very small woodlots (re: Marie Winn’s) • Follow-up to confirm some localities needed
  • 29. Things People Liked …. • Being outside at night looking at something they thought they knew, but really didn’t • Sending in data with their cell phone • Being part of an event • Learning about crickets and katydids in general
  • 30. Dociostaurus (Dociostaurus) maroccanus (Thunberg, 1815) • Prognathogryllus inexspectatus Perkins, 1899 • Trigonidium mahina Otte, 1994 • Gomphocerus plebejus Stål, 1861 • Prognathogryllus kahea Otte, 1994 • Trigonidium makani Otte, 1994 • Gryllidae • Prognathogryllus kahili Otte, 1994 • Trigonidium makanina Otte, 1994 • Anele ulia Otte, Carvalho & Shaw, 2003 • Prognathogryllus kipahulu Otte, 1994 • Trigonidium makapala Otte, 1994 • Caconemobius albus Otte, 1994 • Prognathogryllus kohala Otte, 1994 • Trigonidium makau Otte, 1994 • Caconemobius anahulu Otte, 1994 • Prognathogryllus kukui Otte, 1994 • Trigonidium malanai Otte, 1994 • Caconemobius fori Gurney & Rentz, 1978 • Prognathogryllus makai Otte, 1994 • Trigonidium mana Otte, 1994 • Caconemobius howarthi Gurney & Rentz, 1978 • Prognathogryllus makakapua Otte, 1994 • Trigonidium manuka Carvalho & Otte, 2006 • Caconemobius nihoensis Otte, 1994 • Prognathogryllus mauka Otte, 1994 • Trigonidium mauiensis Otte, 1994 • Caconemobius paralbus Otte, 1994 • Prognathogryllus oahuensis Perkins, 1899 • Trigonidium mauka Otte, 1994 • Caconemobius sandwichensis Otte, 1994 • Prognathogryllus olympus Otte, 1994 • Trigonidium mokuleia Carvalho & Otte, 2006 • Caconemobius uuku Otte, 1994 • Prognathogryllus opua Otte, 1994 • Trigonidium molokaiense (Perkins, 1899) • Caconemobius varius Gurney & Rentz, 1978 • Prognathogryllus parakahili Otte, 1994 • Trigonidium nani Otte, 1994 • Laupala cerasina Otte, 1994 • Prognathogryllus parakukui Otte, 1994 • Trigonidium napua Carvalho & Otte, 2006 • Laupala eukolea Otte, 1994 • Prognathogryllus pararobustus Otte, 1994 • Trigonidium nele Otte, 1994 • Laupala eupacifica Otte, 1994 • Prognathogryllus pihea Otte, 1994 • Trigonidium neogrande Otte, 1994 • Laupala fugax Otte, 1994 • Prognathogryllus puna Otte, 1994 • Trigonidium neokukui Otte, 1994 • Laupala hapapa Otte, 1994 • Prognathogryllus robustus Perkins, 1899 • Trigonidium neovarians Otte, 1994 • Laupala hualalai Otte, 1994 • Prognathogryllus spadix Otte, 1994 • Trigonidium novagintum Otte, 1994 • Laupala kai Otte, 1994 • Prognathogryllus stridulans (Perkins, 1899) • Trigonidium novena Otte, 1994 • Laupala kanaele Otte, 1994 • Prognathogryllus victoriae Otte, 1994 • Trigonidium octavum Otte, 1994 • Laupala kauaiensis Otte, 1994 • Prognathogryllus waikemoi Otte, 1994 • Trigonidium octonalis Otte, 1994 • Laupala kohalensis Otte, 1994 • Prognathogryllus weli Otte, 1994 • Trigonidium ohaka Otte, 1994 • Laupala kokeensis Otte, 1994 • Prolaupala kaalensis Otte, 1994 • Trigonidium ola Otte, 1994 • Laupala kolea Otte, 1994 • Prolaupala kukui Otte, 1994 • Trigonidium olomea Otte, 1994 • Laupala koloa Otte, 1994 • Prolaupala perkinsi Otte, 1994 • Trigonidium ookala Otte, 1994 • Laupala kona Otte, 1994 • Thaumatogryllus cavicola Gurney & Rentz, 1978 • Trigonidium opua Otte, 1994 • Laupala lanaiensis Otte, 1994 • Thaumatogryllus conanti Otte, 1994 • Trigonidium pahiwa Otte, 1994 • Laupala makaio Shaw, 2000 • Thaumatogryllus mauiensis Otte, 1994 • Trigonidium palai Otte, 1994 • Laupala makaweli Otte, 1994 • Thaumatogryllus variegatus Perkins, 1899 • Trigonidium paramana Otte, 1994 • Laupala media Otte, 1994 • Trigonidium acuste Otte, 1994 • Trigonidium paranoe Otte, 1994 • Laupala mediaspisa Otte, 1994 • Trigonidium ahiu Otte, 1994 • Trigonidium paranoho Otte, 1994 • Laupala melewiki Shaw, 2000 • Trigonidium aka Otte, 1994 • Trigonidium paraspilos Otte, 1994 • Laupala molokaiensis Otte, 1994 • Trigonidium akaka Otte, 1994 • Trigonidium paroctonalis Otte, 1994 • Laupala neospisa Otte, 1994 • Trigonidium alina Otte, 1994 • Trigonidium pavida Otte, 1994 • Laupala nigra Otte, 1994 • Trigonidium alternatum (Perkins, 1899) • Trigonidium pipili Otte, 1994 • Laupala nui Otte, 1994 • Trigonidium anoe Otte, 1994 • Trigonidium pololu Otte, 1994 • Laupala oahuensis Otte, 1994 • Trigonidium atroferrugineum (Brunner von Wattenwyl, 1895) • Trigonidium proalina Otte, 1994 • Laupala olohena Otte, 1994 • Trigonidium attenuatum (Perkins, 1899) • Trigonidium procrustum Otte, 1994 • Laupala pacifica Otte, 1994 • Trigonidium awawa Otte, 1994 • Trigonidium promana Otte, 1994 • Laupala paranigra Otte, 1994 • Trigonidium awiwi Otte, 1994 • Trigonidium pseudokua Otte, 1994 • Laupala parapacifica Otte, 1994 • Trigonidium crepitans (Perkins, 1899) • Trigonidium pseudoli Otte, 1994 • Laupala paraprosea Otte, 1994 • Trigonidium debile (Perkins, 1899) • Trigonidium pseudonoe Otte, 1994 • Laupala prosea Otte, 1994 • Trigonidium eumeles Otte, 1994 • Trigonidium pudicum Otte, 1994 • Laupala pruna Otte, 1994 • Trigonidium excultatum Otte, 1994 • Trigonidium puiwa Otte, 1994 • Laupala spisa Otte, 1994 • Trigonidium exiguum Perkins, 1899 • Trigonidium puukani Otte, 1994 • Laupala tantalis Otte, 1994 • Trigonidium exuberans Otte, 1994 • Trigonidium robustum Perkins, 1899 • Laupala vespertina Otte, 1994 • Trigonidium filicum (Perkins, 1899) • Trigonidium roseum (Perkins, 1899) • Laupala waikemoi Otte, 1994 • Trigonidium flectens Otte, 1994 • Trigonidium saltator (Perkins, 1899) • Laupala wailua Otte, 1994 • Trigonidium fortuitum Otte, 1994 • Trigonidium septimum Otte, 1994 • Leptogryllus apicalis Perkins, 1910 • Trigonidium freycinetiae (Perkins, 1899) • Trigonidium sextum Otte, 1994 • Leptogryllus deceptor Perkins, 1910 • Trigonidium fritinnium Otte, 1994 • Trigonidium sibilans Otte, 1994 • Leptogryllus elongatus Perkins, 1899 • Trigonidium grande (Perkins, 1899) • Trigonidium silvicola Otte, 1994 • Leptogryllus forficularis (Brunner von Wattenwyl, 1895) • Trigonidium haawina Otte, 1994 • Trigonidium spilos Otte, 1994 • Leptogryllus fusconotatus Perkins, 1899 • Trigonidium halulu Otte, 1994 • Trigonidium subroseum (Perkins, 1899) • Leptogryllus hanaula Otte, 1994 • Trigonidium hamakua Otte, 1994 • Trigonidium sylvaticum Otte, 1994 • Leptogryllus haupu Otte, 1994 • Trigonidium hamumu Otte, 1994 • Trigonidium triens Otte, 1994 • Leptogryllus kaala Otte, 1994 • Trigonidium hapapa Otte, 1994 • Trigonidium ua Otte, 1994 • Leptogryllus kainalu Otte, 1994 • Trigonidium haupu Otte, 1994 • Trigonidium ulaino Otte, 1994 • Leptogryllus kauaiensis Perkins, 1899 • Trigonidium hehelo Otte, 1994 • Trigonidium varians (Perkins, 1899) • Leptogryllus kawela Otte, 1994 • Trigonidium hoahoa Otte, 1994 • Trigonidium venatum Otte, 1994 • Leptogryllus kipahulu Otte, 1994 • Trigonidium hoku Otte, 1994 • Trigonidium virens Otte, 1994 • Leptogryllus kohala Otte, 1994 • Trigonidium holomua Otte, 1994 • Trigonidium viridiscens (Perkins, 1899) • Leptogryllus lanaiensis Otte, 1994 • Trigonidium hopo Otte, 1994 • Trigonidium wahoi Otte, 1994 • Leptogryllus luteus Otte, 1994 • Trigonidium huapala Otte, 1994 • Trigonidium wai Otte, 1994 • Leptogryllus mauiensis Otte, 1994 • Trigonidium hyperkona Otte, 1994 • Trigonidium waialina Otte, 1994 • Leptogryllus mauumae Otte, 1994 • Trigonidium iao Otte, 1994 • Trigonidium waikua Otte, 1994 • Leptogryllus molokai Otte, 1994 • Trigonidium ignavum Otte, 1994 • Trigonidium waimea Otte, 1994 • Leptogryllus montgomeri Otte, 1994 • Trigonidium illex Otte, 1994 • Trigonidium waipuna Otte, 1994 • Leptogryllus nigrolineatus (Perkins, 1899) • Trigonidium imitans Otte, 1994 • Trigonidium wiki Otte, 1994 • Leptogryllus nigromaculatus Perkins, 1899 • Trigonidium improbum Otte, 1994 • Myrmecophilidae • Leptogryllus oahuensis Otte, 1994 • Trigonidium incongruum Otte, 1994 • Myrmecophilus (Myrmecophilus) quadrispina Perkins, 1899
  • 31. Sam Droege – sdroege@usgs.gov