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UNIT 5: SUPPORT AND TRANSPORT
SYSTEMS IN PLANTS
(CAMPBELL & REECE (2010) CHAPTER 35 &36)
1. ANATOMY OF
DICOTYLEDONOUS PLANTS
• Plants, like multicellular animals, have
organs composed of different tissues,
which in turn are composed of cells
• Three basic organs evolved: roots, stems,
and leaves
• They are organized into a root system and
a shoot system
ANATOMY OF
DICOTYLEDONOUS PLANTS
Dermal, Vascular, and Ground
Tissues
• Each plant organ has dermal, vascular,
and ground tissues
Dermal, Vascular, and Ground
Tissues
Dermal tissue
• In nonwoody plants, the dermal tissue
system consists of the epidermis
• A waxy coating called the cuticle helps
prevent water loss from the epidermis
• In woody plants, protective tissues called
periderm replace the epidermis in older
regions of stems and roots
• Trichomes are outgrowths of the shoot
epidermis and can help with insect defence.
Vascular Tissues
• The vascular tissue system carries out
long-distance transport of materials
between roots and shoots
• The two vascular tissues are xylem and
phloem
• Xylem conveys water and dissolved
minerals upward from roots into the shoots
• Phloem transports organic nutrients from
where they are made to where they are
needed
Ground Tissues
• Tissues that are neither dermal nor
vascular are the ground tissue system
• Ground tissue internal to the vascular
tissue is pith; ground tissue external to the
vascular tissue is cortex
• Ground tissue includes cells specialized for
storage, photosynthesis, and support
CROSS SECTION THROUGH A
DICOT ROOT
CROSS SECTION THROUGH A
DICOT STEM
CROSS SECTION THROUGH A
DICOT LEAF
2. SECONDARY GROWTH
(Chapter 35 p.751-754)
• Secondary growth occurs in
stems and roots of woody plants
but rarely in leaves
• The secondary plant body
consists of the tissues produced
by the vascular cambium and
cork cambium
a. The Vascular Cambium and
Secondary Vascular Tissue
• The vascular cambium is a cylinder of meristematic
cells one cell layer thick
• It develops from undifferentiated parenchyma cells
• Secondary xylem accumulates as wood, and
consists of tracheids, vessel elements and fibers.
• Early wood, formed in the spring, has thin cell walls
to maximize water delivery.
• Late wood, formed in late summer, has thick-
walled cells and contributes more to stem support.
a. The Vascular Cambium and
Secondary Vascular Tissue
• Tree rings are visible where late and early wood
meet, and can be used to estimate a tree’s age
• Dendrochronology is the analysis of tree ring
growth patterns, and can be used to study past
climate change.
• As a tree or woody shrub ages, the older layers of
secondary xylem, the heartwood, no longer
transport water and minerals.
• The outer layers, known as sapwood, still
transport materials through the xylem.
a. The Vascular Cambium and
Secondary Vascular Tissue
3. UPTAKE OF WATER AND
MINERALS INTO THE ROOTS
• The plasma membrane directly controls the
traffic of molecules into and out of the cell.
• In most plant tissues, the cell wall and cytosol are
continuous from cell to cell.
• The cytoplasmic continuum is called the
symplast.
• The cytoplasm of neighboring cells is connected
by channels called plasmodesmata.
• The apoplast is the continuum of cell walls and
extracellular spaces.
Water and minerals can travel through
a plant by three routes:
Transmembrane route: out of one
cell, across a cell wall, and into
another cell.
Symplastic route: via the continuum
of cytosol.
Apoplastic route: via the cell walls
and extracellular spaces.
Water and minerals can travel through
a plant by three routes:
Pathway 1: Transmembrane route
Water and minerals move from the soil (high
WP) through the cell wall, plasmamembrane
and into the cytoplasm of the roothair.
Through the cell wall, plasma membrane
and into the cytoplasm of the cortex cells.
Through the cell wall, plasma membrane
and into the cytoplasm of the endodermis
Through the cell wall, plasma membrane
and into the cytoplasm of the pericycle.
Through the cell wall, plasma membrane
and into the xylem of the plant.
Pathway 2: symplastic route
 Water and minerals flow from the high
WP in the soil, through the cell wall,
plasma membrane and into the
cytoplasm of the roothair.
 Through the plasmodesmata linking the
cytoplasm of the cells into the cytoplasm
of the cortex cells.
 Flowing through the plasmodesmata
into the cytoplasma of the endodermis
cells and pericycle and into the xylem.
Pathway 3: Apoplastic route
 During the Apoplastic route the water and
minerals travel along the cell walls of the
roothairs, cortical cells, and endodermis, but
the endodermis has Casparian strips that
prevent further movement of water, therefore
water enter the cytoplasm of the endodermal
cells and then take the symplastic route
(through cytoplasm and plasmodesmata that
link cytoplasm of cells) into the xylem of the
root.
 D:Chapter_36A_PowerPoint_Lectures36_Lecture_Presentation36_12TransportIn
Roots_A.html
Pathway 3: apoplastic route
Transport of minerals across an
endodermal plasma membrane
 Minerals follow the same routes as water when it moves
through the roothair and cortex cells.
 But when it moves through the endodermal plasma
membrane it does it in the following way:
 An ATP driven pump removes hydrogen ions from the
cell.
 This establishes an electrochemical gradient that
allows potassium ions and other positively charged
ions to cross the membrane via a channel protein.
 Negatively charged mineral ions can cross the
membrane by way of a carrier when they “hitch a
ride” with hydrogen ions, which are diffusing down
their concentration gradient.
Transport of minerals across an
endodermal plasma membrane
3. TRANSPORT OF WATER AND
MINERALS TO THE LEAVES
What makes the upward movement of water
in the xylem of the stems and leaves possible?
 Root pressure
 Cohesion-adhesion-tension
 Transpiration
• Water entering root cells creates a positive
pressure called root pressure.
• It occurs at night and tends to push xylem
sap upwards.
I) Root pressure
 Upward movement of water requiring no
energy.
 Cohesion- Tendency of watermolecules to
cling together and form a continuous water
column.
 Adhesion- Tendency of water molecules to
cling to the sides of the container (xylem)
they are in. It gives the water column extra
strength and prevents it from slipping back
II) Cohesion-adhesion-tension
model
II) Cohesion-adhesion-tension
model
 Transpiration is the loss of water vapour through
the stomata of the leaf.
 The water molecules that evaporate from the cells
into the intercellular airspaces are replaced by
other water molecules from the leaf xylem.
 Because the water molecules are cohesive,
transpiration exerts a pulling force, or tension, that
draws the water column through the xylem to
replace the water lost by the stomata.
 This is also a mechanism by which minerals are
transported throughout the plant body.
 D:Chapter_36A_PowerPoint_Lectures36_Lecture_Presentation36_15Transpiration_A.h
tml
III) Transpiration
High temperature – faster transpiration
High light intensity – faster transpiration
Windy – faster transpiration
High humidity – slower transpiration
D:Chapter_36A_PowerPoint_Lectures36_Lecture_Presentation36_15WaterTransportPlants_
A.html
Environmental factors that influence
the speed of transpiration
Environmental factors that influence
the speed of transpiration
What is guttation and what causes it?
 Root pressure is responsible for guttation.
 Guttation is when drops of water are
forced out of the vein endings along the
edges of leaves called hydathodes.
4. TRANSLOCATION OF SUBSTANCES FROM
THE LEAVES TO THE REST OF THE PLANT
(Chapter 36 p. 779-780)
 The products of photosynthesis are
transported through phloem by the
process of translocation.
 Phloem sap is an aqueous solution that is
high in sucrose.
 It travels from a sugar source to a sugar
sink
 A sugar source is an organ that is a net
producer of sugar, such as mature leaves
 A sugar sink is an organ that is a net consumer
or storer of sugar
 Sugar must be loaded into sieve-tube elements
of the phloem.
 Sugar then moves by symplastic or both
symplastic and apoplastic pathways.
 At the sink, sugar molecules diffuse from the
phloem to sink tissues and are followed by
water.
 D:Chapter_36A_PowerPoint_Lectures36_Lecture_Presentation36_20PhloemTranslocSpring_A.html
 D:Chapter_36A_PowerPoint_Lectures36_Lecture_Presentation36_20PhloemTranslocSummer_A.html
TRANSLOCATION OF SUBSTANCES FROM THE
LEAVES TO THE REST OF THE PLANT

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Ls2 afet unit 5 support and transport in plants

  • 1. UNIT 5: SUPPORT AND TRANSPORT SYSTEMS IN PLANTS (CAMPBELL & REECE (2010) CHAPTER 35 &36)
  • 2. 1. ANATOMY OF DICOTYLEDONOUS PLANTS • Plants, like multicellular animals, have organs composed of different tissues, which in turn are composed of cells • Three basic organs evolved: roots, stems, and leaves • They are organized into a root system and a shoot system
  • 4. Dermal, Vascular, and Ground Tissues • Each plant organ has dermal, vascular, and ground tissues
  • 5. Dermal, Vascular, and Ground Tissues
  • 6. Dermal tissue • In nonwoody plants, the dermal tissue system consists of the epidermis • A waxy coating called the cuticle helps prevent water loss from the epidermis • In woody plants, protective tissues called periderm replace the epidermis in older regions of stems and roots • Trichomes are outgrowths of the shoot epidermis and can help with insect defence.
  • 7. Vascular Tissues • The vascular tissue system carries out long-distance transport of materials between roots and shoots • The two vascular tissues are xylem and phloem • Xylem conveys water and dissolved minerals upward from roots into the shoots • Phloem transports organic nutrients from where they are made to where they are needed
  • 8. Ground Tissues • Tissues that are neither dermal nor vascular are the ground tissue system • Ground tissue internal to the vascular tissue is pith; ground tissue external to the vascular tissue is cortex • Ground tissue includes cells specialized for storage, photosynthesis, and support
  • 9. CROSS SECTION THROUGH A DICOT ROOT
  • 10. CROSS SECTION THROUGH A DICOT STEM
  • 11. CROSS SECTION THROUGH A DICOT LEAF
  • 12.
  • 13. 2. SECONDARY GROWTH (Chapter 35 p.751-754) • Secondary growth occurs in stems and roots of woody plants but rarely in leaves • The secondary plant body consists of the tissues produced by the vascular cambium and cork cambium
  • 14.
  • 15.
  • 16.
  • 17. a. The Vascular Cambium and Secondary Vascular Tissue • The vascular cambium is a cylinder of meristematic cells one cell layer thick • It develops from undifferentiated parenchyma cells • Secondary xylem accumulates as wood, and consists of tracheids, vessel elements and fibers. • Early wood, formed in the spring, has thin cell walls to maximize water delivery. • Late wood, formed in late summer, has thick- walled cells and contributes more to stem support.
  • 18. a. The Vascular Cambium and Secondary Vascular Tissue • Tree rings are visible where late and early wood meet, and can be used to estimate a tree’s age • Dendrochronology is the analysis of tree ring growth patterns, and can be used to study past climate change. • As a tree or woody shrub ages, the older layers of secondary xylem, the heartwood, no longer transport water and minerals. • The outer layers, known as sapwood, still transport materials through the xylem.
  • 19. a. The Vascular Cambium and Secondary Vascular Tissue
  • 20. 3. UPTAKE OF WATER AND MINERALS INTO THE ROOTS • The plasma membrane directly controls the traffic of molecules into and out of the cell. • In most plant tissues, the cell wall and cytosol are continuous from cell to cell. • The cytoplasmic continuum is called the symplast. • The cytoplasm of neighboring cells is connected by channels called plasmodesmata. • The apoplast is the continuum of cell walls and extracellular spaces.
  • 21. Water and minerals can travel through a plant by three routes: Transmembrane route: out of one cell, across a cell wall, and into another cell. Symplastic route: via the continuum of cytosol. Apoplastic route: via the cell walls and extracellular spaces.
  • 22. Water and minerals can travel through a plant by three routes:
  • 23. Pathway 1: Transmembrane route Water and minerals move from the soil (high WP) through the cell wall, plasmamembrane and into the cytoplasm of the roothair. Through the cell wall, plasma membrane and into the cytoplasm of the cortex cells. Through the cell wall, plasma membrane and into the cytoplasm of the endodermis Through the cell wall, plasma membrane and into the cytoplasm of the pericycle. Through the cell wall, plasma membrane and into the xylem of the plant.
  • 24. Pathway 2: symplastic route  Water and minerals flow from the high WP in the soil, through the cell wall, plasma membrane and into the cytoplasm of the roothair.  Through the plasmodesmata linking the cytoplasm of the cells into the cytoplasm of the cortex cells.  Flowing through the plasmodesmata into the cytoplasma of the endodermis cells and pericycle and into the xylem.
  • 25. Pathway 3: Apoplastic route  During the Apoplastic route the water and minerals travel along the cell walls of the roothairs, cortical cells, and endodermis, but the endodermis has Casparian strips that prevent further movement of water, therefore water enter the cytoplasm of the endodermal cells and then take the symplastic route (through cytoplasm and plasmodesmata that link cytoplasm of cells) into the xylem of the root.  D:Chapter_36A_PowerPoint_Lectures36_Lecture_Presentation36_12TransportIn Roots_A.html
  • 27. Transport of minerals across an endodermal plasma membrane  Minerals follow the same routes as water when it moves through the roothair and cortex cells.  But when it moves through the endodermal plasma membrane it does it in the following way:  An ATP driven pump removes hydrogen ions from the cell.  This establishes an electrochemical gradient that allows potassium ions and other positively charged ions to cross the membrane via a channel protein.  Negatively charged mineral ions can cross the membrane by way of a carrier when they “hitch a ride” with hydrogen ions, which are diffusing down their concentration gradient.
  • 28. Transport of minerals across an endodermal plasma membrane
  • 29. 3. TRANSPORT OF WATER AND MINERALS TO THE LEAVES What makes the upward movement of water in the xylem of the stems and leaves possible?  Root pressure  Cohesion-adhesion-tension  Transpiration
  • 30. • Water entering root cells creates a positive pressure called root pressure. • It occurs at night and tends to push xylem sap upwards. I) Root pressure
  • 31.  Upward movement of water requiring no energy.  Cohesion- Tendency of watermolecules to cling together and form a continuous water column.  Adhesion- Tendency of water molecules to cling to the sides of the container (xylem) they are in. It gives the water column extra strength and prevents it from slipping back II) Cohesion-adhesion-tension model
  • 33.
  • 34.  Transpiration is the loss of water vapour through the stomata of the leaf.  The water molecules that evaporate from the cells into the intercellular airspaces are replaced by other water molecules from the leaf xylem.  Because the water molecules are cohesive, transpiration exerts a pulling force, or tension, that draws the water column through the xylem to replace the water lost by the stomata.  This is also a mechanism by which minerals are transported throughout the plant body.  D:Chapter_36A_PowerPoint_Lectures36_Lecture_Presentation36_15Transpiration_A.h tml III) Transpiration
  • 35. High temperature – faster transpiration High light intensity – faster transpiration Windy – faster transpiration High humidity – slower transpiration D:Chapter_36A_PowerPoint_Lectures36_Lecture_Presentation36_15WaterTransportPlants_ A.html Environmental factors that influence the speed of transpiration
  • 36. Environmental factors that influence the speed of transpiration
  • 37. What is guttation and what causes it?  Root pressure is responsible for guttation.  Guttation is when drops of water are forced out of the vein endings along the edges of leaves called hydathodes.
  • 38. 4. TRANSLOCATION OF SUBSTANCES FROM THE LEAVES TO THE REST OF THE PLANT (Chapter 36 p. 779-780)  The products of photosynthesis are transported through phloem by the process of translocation.  Phloem sap is an aqueous solution that is high in sucrose.  It travels from a sugar source to a sugar sink
  • 39.  A sugar source is an organ that is a net producer of sugar, such as mature leaves  A sugar sink is an organ that is a net consumer or storer of sugar  Sugar must be loaded into sieve-tube elements of the phloem.  Sugar then moves by symplastic or both symplastic and apoplastic pathways.  At the sink, sugar molecules diffuse from the phloem to sink tissues and are followed by water.  D:Chapter_36A_PowerPoint_Lectures36_Lecture_Presentation36_20PhloemTranslocSpring_A.html  D:Chapter_36A_PowerPoint_Lectures36_Lecture_Presentation36_20PhloemTranslocSummer_A.html
  • 40. TRANSLOCATION OF SUBSTANCES FROM THE LEAVES TO THE REST OF THE PLANT

Editor's Notes

  1. Ground tissue of roots, consisting mostly of parenchyma cells, fills the cortexThe innermost layer of the cortex is called the endodermis – a cylinder 1 cell thick that forms the boundary with the vascular system (selective barier).Pericycle – outermost cell layer in vascular cylinder adjacent to & just inside endodermis – from which lateral roots ariseLateral roots cannot originate near the roots surface because its vascular system must be continuous with the vascular cylinder at the center of the established root.Casparian strip – waxy material impervious to water & dissolved minerals
  2. Ground tissue consists mostly of parenchyma butcollenchyma just beneath the epidermis provides strength for stems.Sclerenchyma cells, especially fiber cells, also provide support in those parts of the stems that are no longer elongating.Vascular cambium (lateral meristem) – cylinder of dividing cells for growth in thickness – adds layers of vascular tissue called secondary xylem and secondary phloem
  3. Ground tissue of a leaf, a region called the mesophyll is sandwiched between the upper and lower epidermal layers.Mesophyll consists mainly of parenchyma specialised for photosynthesis, including pallisade and spongy mesophyll.
  4. Vascular cambium and cork cambium = lateral meristems for secondary growth (thickness)Vascular cambium adds layers of vascular tissue: secondary xylem (wood) & secondary phloemCork cambium (adds secondary dermal tissue)replaces the epidermis with the thicker, tougher periderm.
  5. Primary growth from activity of apical meristem nears completion – vascular cambium forms.Secondary growth begins – stem thickens as vascular cambium forms secondary xylem to the inside and secondary phloem to the outside.Some initials give rise to vascular rays – connect the secondary xylem with the secondary phloem, move water and nutrients, store carbohydrates and aid in wound repair.Secondary phloem can’t keep pace as cells no longer divide – as a result these tissues & epidermis will ruptureCork cambium develops – produces cork cells which replace epidermis
  6. Cytosol - the semi-fluid portion of the cytoplasm