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Crop plant reproduction




                          ALBIO9700/2006JK
The structural features of a wind
                 pollinated plant
Characteristic    Meadow fescue (wind pollinated)

Petals            no petals (small, inconspicuous, often green)
                  no nectaries
                  no scent
                  no landing platform
Flowers           borne at the end of long stalks, held well above the
                  foliage
                  flowers can appear before the leaves
Stamens           hang outside flower to catch winds
                  stamens and anthers swing freely in air currents

Pollen            light, small, smooth surfaced
                  relatively large amount produced
Stigma            large, feathery and hangs outside, therefore traps pollen
                  more easily
                                                                  ALBIO9700/2006JK
Wind pollination




Meadow fescue (Festuca pratensis)
                                    ALBIO9700/2006JK
The genetic outcomes of self and
         cross pollination
            Self pollination                       Cross pollination


Decreased genetic variation/increased   Increased genetic variation/decreased
genetic uniformity                      genetic uniformity

Increased homozygosity/decreased        Increased heterozygosity/decreased
heterozygosity                          homozygosity

Harmful recessive characteristics more Harmful recessive characteristics less
likely to be expressed                 likely to be expressed

Reduction in gene pool                  Maintenance of gene pool


Inbreeding depression reduces fitness   Gives hybrid vigour (= heterosis =
                                        outbreeding enhancement), so fitness
                                        maintained

                                                                       ALBIO9700/2006JK
Self- vs cross-pollination
• Self-pollination more reliable
• Self-pollination advantageous in harsh
  environments
• Self-pollination results in self-fertilisation
  (inbreeding/genetic disadvantage)

• Cross-pollination is less reliable, more wasteful
  of pollen
• Cross-pollination has genetic advantages
  (producing variation/outbreeding)

                                                   ALBIO9700/2006JK
The structure of the fruit in maize
and the function of the endosperm
• The individual fruit of maize is a dry fruit
  (caryopsis) and contains a single seed
• Seed contains 2 structures:
   – Germ: from which new plant will develop
      • Consists of a miniature plant axis to which is attached ~5
        embryonic leaves and a radicle
      • Source of maize ‘vegetable oil’
   – Endosperm: a store of nutrients which will be made
     available to the germinating seedling until it has
     established sufficient leaf area to photosynthesise
      • Takes up about 2/3 of the volume of the seed and accounts
        for around 86% of its dry weight
      • Principle component – starch and 10% protein (gluten)
      • To provide the nutrition required by the germinating seedling
      • Whole, ground maize meal has 1500 kJ per 100g
                                                               ALBIO9700/2006JK
Corn grain
•   Pericarp (fruit wall protecting corn kernel)
•   Aleurone layer (produces a-amylase, an
    enzyme that breaks down starch and thus
    mobilizes energy for germination)
•   Endosperm (rich in starch which serves as the
    energy source for the germinating seed and
    seedling)
•   Cotyledon (does not serve as the storage
    tissue of the seed )
•   Coleoptile (acts as a protective sheath
    surrounding young leaves and apical meristem
    of a plumule)
•   Plumule (embryonic plant)
•   Apical meristem
•   Radical (embryonic root)
•   Coleorhiza (seals and prevents pathogens
    from entering the kernel through the rupture
    made by the elongating radicle during
    germination)
•   Black layer (once formed, it indicates
    physiological maturity)
                                       ALBIO9700/2006JK
The significance of the inclusion of
 cereal grains in the human diet
• Nutritional value of cereal grains:
   – 70-80% carbohydrates (starch) – important source of
     energy
   – Source of protein (6-14%) – millets, rice, maize; rye,
     barley; wheat, oats – low in lysine
   – Low in fat (2-4%; oats 7.5%) – high in essential fatty
     acids (linoleic acid) – fat found in germ of grain
   – Vitamins – B groups vitamins (thiamin, riboflavin,
     niacin, B6, folic acid, biotin and pantothenic acid) and
     fat soluble vitamin E – deficient in vitamins A, D and
     C
   – Minerals – potassium, calcium, magnesium,
     phosphorus, iron and zinc
   – Fibre – wholegrain meal or flour contain much more
     fibre than grains which have been processed and
     refined
                                                       ALBIO9700/2006JK
ALBIO9700/2006JK

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01 Crop Plant Reproduction

  • 1. Crop plant reproduction ALBIO9700/2006JK
  • 2. The structural features of a wind pollinated plant Characteristic Meadow fescue (wind pollinated) Petals no petals (small, inconspicuous, often green) no nectaries no scent no landing platform Flowers borne at the end of long stalks, held well above the foliage flowers can appear before the leaves Stamens hang outside flower to catch winds stamens and anthers swing freely in air currents Pollen light, small, smooth surfaced relatively large amount produced Stigma large, feathery and hangs outside, therefore traps pollen more easily ALBIO9700/2006JK
  • 3. Wind pollination Meadow fescue (Festuca pratensis) ALBIO9700/2006JK
  • 4. The genetic outcomes of self and cross pollination Self pollination Cross pollination Decreased genetic variation/increased Increased genetic variation/decreased genetic uniformity genetic uniformity Increased homozygosity/decreased Increased heterozygosity/decreased heterozygosity homozygosity Harmful recessive characteristics more Harmful recessive characteristics less likely to be expressed likely to be expressed Reduction in gene pool Maintenance of gene pool Inbreeding depression reduces fitness Gives hybrid vigour (= heterosis = outbreeding enhancement), so fitness maintained ALBIO9700/2006JK
  • 5. Self- vs cross-pollination • Self-pollination more reliable • Self-pollination advantageous in harsh environments • Self-pollination results in self-fertilisation (inbreeding/genetic disadvantage) • Cross-pollination is less reliable, more wasteful of pollen • Cross-pollination has genetic advantages (producing variation/outbreeding) ALBIO9700/2006JK
  • 6. The structure of the fruit in maize and the function of the endosperm • The individual fruit of maize is a dry fruit (caryopsis) and contains a single seed • Seed contains 2 structures: – Germ: from which new plant will develop • Consists of a miniature plant axis to which is attached ~5 embryonic leaves and a radicle • Source of maize ‘vegetable oil’ – Endosperm: a store of nutrients which will be made available to the germinating seedling until it has established sufficient leaf area to photosynthesise • Takes up about 2/3 of the volume of the seed and accounts for around 86% of its dry weight • Principle component – starch and 10% protein (gluten) • To provide the nutrition required by the germinating seedling • Whole, ground maize meal has 1500 kJ per 100g ALBIO9700/2006JK
  • 7. Corn grain • Pericarp (fruit wall protecting corn kernel) • Aleurone layer (produces a-amylase, an enzyme that breaks down starch and thus mobilizes energy for germination) • Endosperm (rich in starch which serves as the energy source for the germinating seed and seedling) • Cotyledon (does not serve as the storage tissue of the seed ) • Coleoptile (acts as a protective sheath surrounding young leaves and apical meristem of a plumule) • Plumule (embryonic plant) • Apical meristem • Radical (embryonic root) • Coleorhiza (seals and prevents pathogens from entering the kernel through the rupture made by the elongating radicle during germination) • Black layer (once formed, it indicates physiological maturity) ALBIO9700/2006JK
  • 8. The significance of the inclusion of cereal grains in the human diet • Nutritional value of cereal grains: – 70-80% carbohydrates (starch) – important source of energy – Source of protein (6-14%) – millets, rice, maize; rye, barley; wheat, oats – low in lysine – Low in fat (2-4%; oats 7.5%) – high in essential fatty acids (linoleic acid) – fat found in germ of grain – Vitamins – B groups vitamins (thiamin, riboflavin, niacin, B6, folic acid, biotin and pantothenic acid) and fat soluble vitamin E – deficient in vitamins A, D and C – Minerals – potassium, calcium, magnesium, phosphorus, iron and zinc – Fibre – wholegrain meal or flour contain much more fibre than grains which have been processed and refined ALBIO9700/2006JK